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For stocks grown to a size sufficient to accumulate numerous mutations, the general formulation is μ.

Note that this expression holds only in the absence of selection, which is likely to be weak or absent during the first cycle of infection but may become strong in subsequent cycles.

Each culture is then screened simply for the presence or absence of mutants, and the average number of virus particles per culture is also determined.

On the assumption that mutational events are randomly distributed among replication events and because the number of replication events per culture is close to is a third of the total number of different base substitutions that can be monitored.) In these riboviral systems, only base-pair substitutions are scored.

When rates of spontaneous mutation are expressed per genome per genome replication, different broad groups of organisms display characteristic values (3): roughly 0.2 for retroelements; close to 0.0034 for DNA-based microbes (including both viral and cellular organisms); and roughly 0.01 for higher eukaryotes.

will generally be a small (but not necessarily an integer) number, from one for a single round of infection (high multiplicity of infection), to perhaps four or five for a large population of host cells and a small inoculum.

In addition, the formulation also suggests ways to measure hitherto opaque parameters of viral replication.

The replication of riboviruses whose chromosomes are composed of single-stranded RNA follows a simple scheme (Fig. A cell is infected with one (or more) virus particles.

The number of sites at which mutation could occur was estimated from the observation that five sequenced mutations fell into four sites; assuming a Poisson distribution of mutations among sites, the most probable number of sites is 7.5.

The size of the measles genome is 15,894 bases (14).




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A racist jury acquitted his murderers, Roy Bryant and J. Milam, despite overwhelming evidence, and, to rub salt in the wound, both admitted to killing Till in magazine the next year. The shoe was on the other foot for once and so be it if two white people wound up dead. That's harsh, but that's the historical context of black men dating white women that I unfortunately have to consider when doing the same. Even then, I understood that it was racial, but there was a disconnection from my personal reality. Whatever I learned from the trial was tucked away as something that I should know as a black man, but it didn't have a life-altering impact on my own development. For whatever implications the trial had, that shit also had nothing to do with me. My attraction to them was likely a natural response to my environment. Nobody was trying to assimilate with white people, but sometimes that's just the way things go when you want a better home and better schools for your family.


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Many parsers, for example, return separate text events for numeric character references.